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The plot concerns Lewis Thomas (Paul Walker), a young man who decides to go cross-country to pick up his high school sweetheart, Venna Wilcox (Leelee Sobieski), from college. On his way there, he bails his brother, Fuller Thomas (Steve Zahn), out of jail. Fuller, looking for a good time, decides to ride with Lewis. During their trip to pick up Venna, Fuller coaxes Lewis into playing a cruel joke, through their CB radio, on a truck driver, known only as ‘Rusty Nail’ (voiced by Ted Levine). When the joke takes a deadly turn, Lewis, Venna, and Fuller find themselves being relentlessly stalked by the psychotic truck driver who is looking for more than an apology.

Donnie Darko gets along badly with his family, with his teachers and with his classmates; but he does manage to find a sympathetic friend in Gretchen, who agrees to date him; and he has a compassionate psychiatrist, who discovers hypnosis is the means to unlock hidden secrets. His other companion may not be a true ally. Donnie has a friend named Frank, who is either a large purple bunny, or man in a large purple bunny costume. Either way, Donnie is the only one who can see him. When an engine falls off a plane and destroys his room, Donnie is not there. Both the event, and Donnie’s escape, seem to have been caused by supernatural events. Donnie’s mental illness, if such it is, may never allow him to find out for sure.